College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China; Beijing Yanshan Earth Critical Zone National Research Station, University of Chinese Academy of Sciences, Beijing 101408, China. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China; Beijing Yanshan Earth Critical Zone National Research Station, University of Chinese Academy of Sciences, Beijing 101408, China; State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Chinese Academy of Sciences, Beijing 100101, China. Electronic address: yfwang@ucas.ac.cn. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China. College of Life Sciences, University of Chinese Academy of Sciences, Beijing 100049, China; Beijing Yanshan Earth Critical Zone National Research Station, University of Chinese Academy of Sciences, Beijing 101408, China. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China; Beijing Yanshan Earth Critical Zone National Research Station, University of Chinese Academy of Sciences, Beijing 101408, China; Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810001, China; Binzhou Institute of Technology, Weiqiao-UCAS Science and Technology Park, Binzhou 256606, China.